† Background and Aims: Rhamnogalacturonan-II (RG-II) is one of the pectin motifs found in the cellwall of all land plants. It contains sugars such as 2-keto-3-deoxy-D-lyxo-heptulosaric acid (Dha) and 2-keto-3-deoxy-D-manno-octulosonic acid (Kdo), and within thewall RG-II is mostly found as a dimer via a borate diester cross-link.To date, little is knownregarding the biosynthesis of this mot…